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Project Gutenberg's Encyclopedia, vol. 1 ( A - Andropha

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description will here begin with the oesophagus or gullet. 

The oesophagus (Gr. oiso, I will carry, and fagein, to 
eat), a muscular tube lined with mucous membrane, stretches 
from the lower limit of the pharynx, at the level of the 
cricoid cartilage, to the cardiac orifice of the stomach.  
It is about 10 in. long (25 cm.) and half to one inch in 
diameter.  At first it lies in the lower part of the neck, 
then in the thorax, and lastly, for about an inch, in the 
abdomen.  As far as the level of the fourth or fifth thoracic 
vertebra it lies behind the trachea, but when that tube ends, 
it is in close contact with the pericardium, and, at the level 
of the tenth thoracic vertebra, passes through the oesophageal 
opening of the diaphragm (q.v.), accompanied by the two 
vagi nerves, the left being in front of it and the right 
behind.  In the abdomen it lies just behind the left lobe of the 
liver.  Both in the upper and lower parts of its course it lies 
a little to the left of the mid line.  Its mucous membrane is 
thrown into a number of longitudinal pleats to allow stretching. 

The stomach (Gr. stomachos) is an irregularly pear-shaped 
bag, situated in the upper and left part of the abdomen.  
It is somewhat flattened from before backward and so has 
an anterior and posterior surface and an upper and lower 
border.  When moderately distended the thick end of the 
pear or fundus bulges upward and to the left, while 
the narrow end is constricted to form the pylorus, by 
means of which the stomach communicates with the small 
intestine.  The cardiac orifice, where the oesophagus 
enters, is placed about a third of the way along the upper 
border from the left end of the fundus, and, between it and 
the pylorus, the upper border is concave and is known as 
the lesser curvature.  From the cardiac to the pyloric 
orifice, round the lower border, is the greater curvature.  
The stomach has in front of it the liver (see fig. 1), the 
diaphragm and the anterior abdominal wall, while behind it 
are the pancreas, left kidney, left adrenal, spleen, colon and 
mesocolon.  These structures form what is known as the stomach 
chamber.  When the stomach is empty it contracts into a tubular 
organ which is frequently sharply bent, and the transverse 
colon ascends to occupy the vacant part of the stomach chamber. 

The last inch of the stomach before reaching the pylorus is 


From A. Birmingham; Cunningham's Text-Book of Anatomy. 

FIG. 1.--The Abdominal Viscera in situ, as seen when the abdomen 
is laid open and the great omentum removed (drawn to scale from a 
photograph of a male body aged 56, hardened by formalin injections). 

The ribs on the right side are indicated by Roman numerals; 
it will be observed that the eighth costal cartilage 
articulated with the sternum on both sides.  The subcostal, 
intertubercular, and right and left Poupart lines are drawn 
in black, and the mesial plane is indicated by a dotted 
line.  The intercostal muscles and part of the diaphragm 
have been removed, to show the liver and stomach extending up 
beneath the ribs.  The stomach is moderately distended, and 
the intestines are particularly regular in their arrangement. 


usually tubular and is known as the pyloric canal. Before 
reaching this there is a bulging known as the pyloric vestibule 
(see D. J. Cunningham, Tr. R. Soc. of Edinib. vol. xlv. 
pt. 1, No. 2). The pylorus is an oval opening, averaging 
half an inch in its long axis but capable of considerable 
distension; it is formed by a special development of the 
circular muscle layer of the stomach, and during life is 
probably tightly closed.  The mucous membrane of the stomach 
is thrown into pleats or rugae when the organ is not fully 
distended, while between these it has a mammillated appearance. 

Superficial to the mucous coat is a sub-mucous, consisting 
of loose connective tissue, while superficial to this are 
three coats of unstriped muscle, the inner oblique, the middle 
circular and the outer longitudinal.  The peritoneal coat is 
described in the article on the coelom and serous membranes. 

The small intestine is a tube, from 22 to 25 ft. long, 
beginning at the pylorus and ending at the ileo-caecal 
valve; it is divided into duodenum, jejunum and ileum. 

The duodenum is from 9 to 11 in. long and forms a horseshoe 
or C-shaped curve, encircling the head of the pancreas.  It 
differs from the rest of the gut in being retroperitoneal.  
Its first part is horizontal and lies behind the fundus of 
the gall-bladder, passing backward and to the right from the 
pylorus.  The second part runs vertically downward in 
front of the hilum of the right kidney, and into this part 
the pancreatic and bile ducts open.  The third part runs 
horizontally to the left in front of the aorta and vena 
cava, while the fourth part ascends to the left side of 
the second lumbar vertebra, after which it bends sharply 
downward and forward to form the duodeno-jejunal flexure. 

The jejunum forms the upper two-fifths of the rest of 
the small intestine; it, like the ileum, is thrown into 
numerous convolutions and is attached by the mesentery to the 
posterior abdominal wall. (See COELOM AND SEROUS MEMBRANES.) 

The ileum is the remaining three-fifths of the small 
intestine, though there is no absolute point at which the 
one ends and the other begins.  Speaking broadly, the jejunum 
occupies the upper and left part of the abdomen below the 
subcostal plane (see ANATOMY: Superficial and Artistic), 
the ileum the lower and right part.  About 3 ft. from its 
termination a small pouch, known as Meckel's diverticulum, 
is very occasionally found.  At its termination the ileum 
opens into the large intestine at the ileo-caecal valve. 

The caecum is a blind sac occupying the right iliac fossa and 
extending down some two or three inches below the ileo- caecal 
junction.  From its posterior and left surface the vermiform 
appendix protrudes, and usually is directed upward and to the 
left, though it not infrequently hangs down into the true 
pelvis.  This worm-like tube is blind at its end and is 
usually 3 or 4 in. long, though it has been seen as long as 10. 
in.  Its internal opening into the caecum is about 1 in. below 
that of the ileum.  On transverse section it is seen to be 
composed of (1) an external muscular coat, (2) a submucous 
coat, (3) a mass of lymphoid tissue, which appears after 
birth, and (4) mucous membrane.  In many cases its lumen is 
wholly or partly obliterated, though this is probably due to 
disease (see R. Berry and L. Lack, Journ.  Anat. & Phys. 
vol.  H. p. 247).  Guarding the opening of the ileum into 
the caecum is the ileo-caecal valve, which consists of two 
cusps projecting into the caecum; of these the upper forms a 
horizontal shelf, while the lower slopes up to it obliquely.  
Complete absence of the valve has been noticed, and in one 
such case the writer found that no abdominal inconvenience had 
been recorded during life.  The caecum is usually completely 
covered by peritoneum, three special pouches of which are 
often found in its neighbourhood; of these the ileo-colic is 
just above the point of junction of the ileum and caecum, the 
ileocaecal just below that point, while the retro-caecal is 
behind the caecum.  At birth the caecum is a cone, the apex 
of which is the appendix; it is bent upon itself to form a 
U, and sometimes this arrangement persists throughout life 
(see C. Toldt, ``Die Formbildung d. menschl.  Blinddarmes,'' 
Sitz. der Wiener Akad. Bd. ciii.  Abteil. 3, p. 41). 

The ascending colon runs up from the caecum at the level of the 
ileo-caecal valve to the hepatic flexure beneath and behind the 
right lobe of the liver; it is about 8 in. long and posteriorly 
is in contact with the abdominal wall and right kidney.  It is 
covered by peritoneum except on its posterior surface (see fig. 1). 

The transverse colon is variable in position, depending largely 
on the distension of the stomach, but usually corresponding 
to the subcostal plane (see ANATOMY: Superficial and 
Artistic).  On the left side of the abdomen it ascends to the 
splenic flexure, which may make an impression on the spleen (see 
DUCTLESS GLANDS), and is bound to the diaphragm opposite the 
eleventh rib by a fold of peritoneum called the phrenico-colic 
ligament.  The peritoneal relations of this part are 
discussed in the article on the coelom and serous membranes. 

The descending colon passes down in front of the left kidney 
and left side of the posterior abdominal wall to the crest 
of the ilium; it is about 6 in. long and is usually empty and 
contracted while the rest of the colon is distended with gas; 
its peritoneal relations are the same as those of the ascending 
colon, but it is more likely to be completely surrounded. 

The iliac colon stretches from the crest of the ilium 
to the inner border of the psoas muscle, lying in the left 
iliac fossa, just above and parallel to Poupart's ligament.  
Like the descending, it is usually uncovered by peritoneum 
on its posterior surface.  It is about 6 in. in length. 

The pelvic colon lies in the true pelvis and forms a 
loop, the two limbs of which are superior and inferior 
while the convexity reaches across to the right side of the 
pelvis.  In the foetus this loop occupies the right 
iliac fossa, but, as the caecum descends and enlarges 
and the pelvis widens, it is usually driven out of this 
region.  The distal end of the loop turns sharply downward 
to reach the third piece of the sacrum where it becomes the 
rectum.  To this pelvic colon Sir F. Treves (Anatomy of 
the Intestinal Canal, London, 1885) has given the name 
of the omega loop.  Formerly the iliac and pelvic colons 
were spoken of as the sigmoid flexure, but Treves and T. 
Jonnesco (Le Colon pelvien pendant la vie intra-uterine, 
Paris, 1892) have pointed out the inapplicability of the 
term, and to the latter author the modern description is due. 

The rectum, according to modern ideas, begins in front of 
the third piece of the sacrum; formerly the last part of the 
O (or omega) loop was described as its first part.  It ends 
in a dilatation or rectal ampulla, which is in contact with 
the back of the prostate in the male and of the vagina in the 
female and is in front of the tip of the coccyx.  The rectum 
is not straight, as its name would imply, but has a concavity 
forward corresponding to that of the sacrum and coccyx. 

When viewed from in front three bends are usually seen, the 
upper and lower of which are sharply concave to the left, 
the middle one to the right.  At the end of the pelvic colon 
the mesocolon ceases, and the rectum is then only covered 
by peritoneum at its sides and in front; lower down the 
lateral covering is gradually reflected off and then only 
the front is covered.  About the junction of the middle and 
lower thirds of the tube the anterior peritoneal covering 
is also reflected off on to the bladder or vagina, forming 
the recto-vesical pouch in the male and the pouch of 
Douglas in the female.  This reflexion is usually about 3 
in. above the anal aperture, but may be a good deal lower. 

The anal canal is the termination of the alimentary tract, 
and runs downward and backward from the lower surface of 
the rectal ampulla between the levatores ani muscles.  It 
is about an inch long and its lateral walls are in contact, 
so that in section it appears as an antero-posterior slit 
(see J. Symington, Journ.  Anat. and Phys. vol. 23, 1888). 

Structure of the Intestine.--The intestine has four coats: 
serous, muscular, submucous and mucous.  The serous or 
peritoneal coat has already been described wherever it is 
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